Biology
Honey bees live in social units called colonies. A honey bee colony
consists of a single queen, who is usually the mother of all other colony
members, ten to thirty thousand semi-sterile female workers and, from
zero to a few thousand males (drones) depending on the time of year.
The immature forms are collectively referred to as brood and include
eggs, larvae, and pupae. Adult workers perform all of the behavioral
tasks associated with colony living. Worker honey bees perform different
tasks as they age, a phenomenon called temporal polyethism. When bees
emerge from their cells as adults they normally clean cells, then as
they age they feed larvae (nursing behavior), process and store food,
secrete wax and construct comb, and guard the entrance. The most pronounced
change in behavior occurs when bees are about three weeks old when they
begin foraging. At this time, they cease performing most tasks within
the nest and usually remain foragers for the rest of their lives. This
age progression is not rigid and many factors such as genotype, colony
social structure, foraging environment, time of year, and pheromones
affect developmental rates and trajectories.
Colonies reproduce through a process of colony budding, commonly referred
to as swarming. As colonies grow to swarming size, factors that
inhibit queen rearing are released and new queens are reared in the presence
of the old queen. Ordinarily, but not necessarily, the old queen leaves
the nest with a group of bees to initiate a new nest. A daughter queen
inherits the valuable parental nest and any remaining adults. In feral,
unmanaged colonies, swarming-related queen rearing begins when colony
size is about 12,000 workers and colonies may swarm when there are about
20,000 workers. Adult population size at swarming is highly variable
and not the singular determinant for swarming. Other factors such as
time of year, adult age distribution, and foraging environment also affect
the timing of swarming. In temperate climates swarming usually takes
place in the spring. In tropical and sub-tropical climates swarming may
take place any time of year. |